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AP Biology Notes

8.5.3 Effects of Competition and Symbiosis on Populations

In community ecology, the interactions between organisms, particularly through competition, predation, and various forms of symbiosis, significantly influence the dynamics of populations. These interactions are fundamental in shaping both individual species and the overall structure of ecological communities.

Competition in Communities

Competition occurs when organisms vie for limited resources such as food, space, or mates. This struggle can significantly impact population dynamics and community structure.

Intraspecific Competition

  • Definition: Competition within the same species.

  • Effects: Can lead to reduced resources availability, lower reproductive rates, and increased mortality.

  • Example: In a deer population, individuals compete for limited forage, impacting growth and survival rates.

Interspecific Competition

  • Definition: Competition between different species.

  • Competitive Exclusion Principle: Describes how two species competing for identical resources cannot coexist indefinitely.

  • Resource Partitioning: Species evolve to use different resources or engage in different behaviors to reduce competition.

  • Example: Different bird species may feed on the same tree but at different heights to reduce competition.

Predation and Population Dynamics

Predation is a biological interaction where one organism, the predator, hunts and feeds on another organism, the prey. This interaction is a critical driver of ecological dynamics.

Predator-Prey Relationships

  • Dynamic Nature: These relationships can lead to oscillations in the population sizes of both predators and prey.

  • Population Control: Predators can keep prey populations in check, preventing overconsumption of resources.

Trophic Cascades

  • Definition: Predators at high trophic levels indirectly affect organisms at lower trophic levels by controlling the population of their prey.

  • Example: The reintroduction of wolves in Yellowstone National Park led to changes in elk populations, which in turn affected plant communities and river systems.

Adaptations for Predation and Defense

  • Predatory Adaptations: Include development of acute senses, speed, stealth, and physical features like claws and teeth.

  • Prey Defenses: Include camouflage, defensive structures, chemical defenses, and behavioral strategies like flocking or schooling.

Symbiosis and Its Impact

Symbiosis refers to a close and long-term biological interaction between two different biological organisms, significantly affecting population dynamics.

Parasitism

  • Characteristics: One organism (the parasite) benefits at the expense of the other (the host).

  • Impact on Host: Can lead to decreased host fitness and survival, potentially influencing host population dynamics.

  • Co-evolution: Hosts and parasites often undergo co-evolution, where changes in one lead to adaptations in the other.

Mutualism

  • Mutual Benefit: Both organisms benefit from the interaction.

  • Enhanced Survival and Growth: Mutualistic relationships can lead to improved resource acquisition, protection from predators, or reproductive advantages.

  • Example: The relationship between coral and zooxanthellae, where coral provides a habitat, and zooxanthellae contribute to the coral's energy needs through photosynthesis.

Commensalism

  • One-Sided Benefit: One organism benefits, and the other is neither helped nor harmed.

  • Examples: An epiphytic plant growing on a tree or birds nesting in trees.

Influence on Community-Level Interactions

Altering Species Composition

  • Selective Pressures: These interactions can act as selective pressures, influencing the traits that are favorable in a community.

  • Species Turnover: Changes in population dynamics due to these interactions can lead to species turnover, affecting community composition.

Influence on Species Diversity

  • Diversity Increase: Predation can reduce the dominance of certain species, allowing less competitive species to thrive.

  • Diversity Decrease: Intense competition or parasitism can decrease species diversity by eliminating weaker species.

Balancing and Regulating Populations

Natural Control Mechanisms

  • Ecological Equilibrium: These interactions help maintain ecological balance, preventing any one species from dominating the ecosystem.

  • Population Checks: Serve as natural checks on population sizes, contributing to the stability of ecosystems.

Indicator of Ecosystem Health

  • Ecosystem Health: The nature and extent of these interactions can indicate the health and stability of an ecosystem.

  • Biodiversity Indicators: High levels of symbiosis and balanced predator-prey dynamics often signify a healthy, diverse ecosystem.

Human Impact on Competition and Symbiosis

Human activities can have profound impacts on these natural interactions, often leading to ecological imbalances.

Invasive Species

  • Competition Increase: Invasive species can outcompete native species, leading to decreased biodiversity.

  • Predator-Prey Imbalance: Introduction of new predators or elimination of native predators can disrupt established predator-prey dynamics.

Habitat Loss

  • Resource Scarcity: Habitat destruction reduces the availability of resources, intensifying competition.

  • Symbiotic Relationship Disruption: Loss of habitat can disrupt important symbiotic relationships, impacting the survival of dependent species.

FAQ

Intraspecific competition can significantly influence the genetic diversity of a species. This competition arises when individuals of the same species compete for limited resources such as food, shelter, and mates. It leads to what is known as 'differential survival and reproduction,' a principle core to the theory of natural selection. Individuals with traits that give them an advantage in competing for these resources are more likely to survive and reproduce. Over time, these advantageous traits become more common within the population, potentially leading to changes in the genetic composition of the species.

This process can enhance genetic diversity if the traits that confer competitive advantages are diverse and change over time, or it can reduce genetic diversity if a specific trait becomes overwhelmingly dominant. For instance, in a plant species competing for sunlight, those with a genetic disposition for taller growth might outcompete shorter plants, leading to a population predominantly composed of taller plants. However, if environmental conditions change, say less sunlight due to canopy closure, plants with traits favoring shade tolerance might then have a competitive edge. This dynamic interaction with the environment ensures that intraspecific competition continually molds the genetic landscape of a species.

Yes, symbiotic relationships can evolve into parasitic relationships over time, a process driven by changes in the ecological environment or genetic mutations. In a symbiotic relationship, both organisms benefit from the interaction. However, if the balance of costs and benefits shifts, one organism might start exploiting the other, leading to a parasitic relationship. This shift can happen if the 'cost' of the relationship becomes too high for one partner, or if 'cheating' behaviors evolve that provide more benefit to one organism at the expense of the other.

For example, in a mutualistic relationship where one organism provides food in exchange for protection, a genetic mutation in the protecting organism might lead it to consume more resources while offering less protection. Over time, if these traits are favored by natural selection, the relationship could become parasitic. Evolutionary changes in one partner can impose selective pressures on the other, potentially leading to a continuous co-evolutionary arms race, where each organism evolves strategies either to exploit the other more effectively or to defend against exploitation. This dynamic nature of biological interactions means that the line between symbiosis and parasitism can be fluid and subject to evolutionary change.

Changes in predator-prey dynamics can have profound effects on the overall biodiversity of an ecosystem. Predators play a crucial role in maintaining the balance of ecosystems by controlling the population of prey species, which in turn affects the populations of other species in the food web. When predator populations decline, this can lead to an overabundance of prey species, which may over-consume primary producers like plants. This imbalance can lead to habitat degradation and a decrease in biodiversity as plant species are lost.

Conversely, if prey populations decline, perhaps due to over-predation or other factors like disease, predators may struggle to find sufficient food. This can lead to a decrease in predator populations and can affect the balance of competitive relationships among prey species, potentially allowing some species to dominate and reduce overall biodiversity. Furthermore, predators often target specific prey species, including those that are particularly competitive or aggressive. By controlling these key species, predators maintain ecological diversity by providing opportunities for less dominant species to thrive. Thus, maintaining healthy predator-prey dynamics is crucial for preserving ecosystem biodiversity.

Interspecific competition is a significant driver of evolutionary change. When species compete for the same limited resources, such as food, water, or habitat, there is a powerful selective pressure on each species to adapt in ways that reduce competition. These adaptations can lead to evolutionary changes that enhance survival and reproductive success in the competitive environment.

One of the primary evolutionary outcomes of interspecific competition is resource partitioning, where competing species evolve to utilize different resources or different aspects of the same resource. For instance, different bird species might evolve to forage at different heights in the same tree or at different times of the day to avoid direct competition. Another outcome is character displacement, where species that compete for similar resources evolve distinct physical or behavioral traits that minimize competition. An example is the Galápagos finches, which evolved different beak shapes to exploit different food sources.

These evolutionary changes driven by interspecific competition can lead to increased niche specialization, reducing direct competition and allowing coexistence. This process enhances the diversity of life forms within an ecosystem, contributing to the overall biodiversity and ecological complexity.

Keystone species have a disproportionately large impact on their ecosystems relative to their abundance, and their presence can significantly influence competition and symbiotic relationships. A keystone species can be a top predator, a mutualistic organism, or any other species that holds a critical role in maintaining the structure of an ecosystem.

When a keystone species is a top predator, it can control the populations of competitive dominant species, thereby reducing competitive exclusion and allowing a greater number of species to coexist. This control can increase species diversity and stabilize the ecosystem. For example, sea otters are keystone predators that control sea urchin populations, which in turn helps maintain kelp forest ecosystems.

In cases where a keystone species is part of a mutualistic relationship, their actions can enhance the survival and success of other species. For instance, certain species of figs and fig wasps are mutually dependent, with each species playing a critical role in the reproductive success of the other. The loss of such a keystone species can disrupt these relationships, leading to cascading effects throughout the ecosystem.

The presence of keystone species often creates a more stable and resilient ecosystem, where competitive pressures are balanced, and symbiotic relationships are maintained, contributing significantly to ecological integrity and biodiversity.

Practice Questions

A biologist studying a grassland ecosystem observes a decline in the population of a native grass species after the introduction of a new grass species from a different continent. The new species is very similar in its requirements for sunlight and nutrients. Which of the following best explains the decline in the native grass species population?

The decline in the native grass species is likely due to interspecific competition with the newly introduced grass species. Interspecific competition occurs when two or more species in the same ecological area compete for the same resources, such as sunlight and nutrients, which are necessary for their growth and survival. In this scenario, the introduction of a similar grass species creates a situation where both species are competing for the same limited resources. The competitive exclusion principle suggests that two species competing for identical resources cannot coexist at constant population values; one species will eventually outcompete the other. The decline in the native grass population indicates that the new species may be more efficient in resource utilization, leading to competitive exclusion of the native species.

In a coral reef ecosystem, small fish known as cleaner wrasse feed on parasites found on the bodies of larger fish. In return, the larger fish do not eat the cleaner wrasse and provide them with a steady food source. Which type of symbiotic relationship does this scenario best exemplify, and how does it affect the population dynamics of both species?

This scenario exemplifies a mutualistic symbiotic relationship, where both the cleaner wrasse and the larger fish benefit from the interaction. The cleaner wrasse gain a reliable food source by feeding on the parasites, while the larger fish benefit from parasite removal, which can improve their health and fitness. This mutualistic relationship likely contributes to the stability of both populations. For the cleaner wrasse, the relationship provides a steady food source and protection from being preyed upon by the larger fish. For the larger fish, having parasites removed can lead to better health, potentially increasing their lifespan and reproductive success. Consequently, this mutualism can contribute to the overall health and biodiversity of the coral reef ecosystem, as it helps maintain balanced population dynamics and encourages species diversity.

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